Bulimulidae
Specimen of Drymaeus poecilus crawling over a piece of tree bark at El Impenetrable National Park of Chaco Province, Argentina
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Suborder:	Helicina
Superfamily:	Orthalicoidea
Family:	Bulimulidae Tryon, 1867
Genera
See text
Diversity[1]
1243 species (including Simpulopsidae and Bothriembryon).

Bulimulidae is a taxonomic family of medium-sized to large, air-breathing, tropical and sub-tropical land snails, terrestrial pulmonate gastropod molluscs in the superfamily Orthalicoidea.^[2]

Distribution of species in the family Bulimulidae includes Ecuador (9 genera)^[3] and other South American countries. Some species also occur in North America.

The family's oldest fossil record dates from the late Cretaceous of Brazil (Itaboraí Basin).^[4][5]

Members of this family have a haploid chromosome number between 26 and 30 (according to the values in this table).^[6]

Shells of species within Bostrycinae have a smooth protoconch and the genital organs feature a relatively long penis sheath (ca. 1/4–1/6 total phallus length).^[7]

Previously, the members of the Orthalicidae were also included in this family, as the subfamily Orthalicinae, and the taxa listed here were placed in their own subfamily, the Bulimulinae.^[8]

Bulimulinae was placed in the family Orthalicidae according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005).^[9]

The subfamily Bulimulinae included 3 tribes:^[9]

• Tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
• Tribe Odontostomini Pilsbry & Vanatta, 1898 - synonym: Tomogeridae Jousseaume, 1877
• Tribe Simpulopsini Schileyko, 1999

Breure et al. (2010)^[1] moved Bothriembryon to Placostylidae (since 2012 as Bothriembryontidae), elevated Bulimulinae to Bulimulidae and Odontostomini to Odontostomidae.^[1]

Breure & Romero (2012)^[10] confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae.^[10] There are three subfamilies within Bulimulidae:^[10]

• subfamily Bulimulinae Tryon, 1867
• subfamily Peltellinae Gray, 1855
• subfamily Bostrycinae Breure, 2012^[10][7]

Genera in the family Bulimulidae include:

subfamily Bulimulinae Tryon, 1887

(synonym: Berendtiinae P. Fischer & Crosse, 1872)

• Anctus E. von Martens, 1860
• Auris Spix, 1827
• Berendtia Crosse & P. Fischer, 1869
• Bocourtia Rochebrune, 1882
• Bulimulus Leach, 1814 - type genus of the subfamily Bulimulinae^[9][3][10]
• Cochlorina Jan, 1830
• Graptostracus Pilsbry, 1939
• Kora Simone, 2012^[11]
• Lopesianus Weyrauch, 1958
• Naesiotus Albers, 1850^{[12][13][3][10]}
• Neopetraeus E. von Martens, 1885
• Newboldius Pilsbry, 1932
• Otostomus H. Beck, 1837
• Oxychona Mörch, 1852
• Protoglyptus Pilsbry, 1897^[14]
• Pseudoxychona Pilsbry, 1930
• Rabdotus Albers, 1850^[10]
• Scutalus Albers, 1850^[15][3][10]
• Spartocentrum Dall, 1895
• Sphaeroconcha Breure, 1978
• Stenostylus Pilsbry, 1898^[13][3][14]
• Suniellus Breure, 1978^[14]

subfamily Peltellinae

• Antidrymaeus Germain, 1907^[16]
• Drymaeus Albers, 1850^[13][3][10]
• Peltella Gray, 1855 - type genus of the subfamily Peltellinae^[10]
• Mesembrinus Albers, 1850^[16]

subfamily Bostrycinae

• Bostryx Troschel, 1847 sensu lato^[3][10]- type genus of the subfamily Bostrycinae^[10]

Unassigned within Bulimulidae

• Bulimus Bruguière, 1789 (temporary name)
• †Itaborahia Maury, 1935^[4]
• † Oreoconus D. W. Taylor in McKenna et al., 1962
• † Palaeobulimulus Parodiz, 1949
• Stapafurdius Simone, 2021
• † Tocobaga Auffenberg, Slapcinsky & Portell, 2015

• Adzharia P. Hesse, 1933: synonym of Bulimulus Leach, 1814 (junior synonym)
• Antidrymaeus Germain, 1907: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
• Atahualpa Strebel, 1910: synonym of Thaumastus E. von Martens, 1860
• Ataxellus Dall, 1912: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented asBostryx Troschel, 1847
• Ataxus Albers, 1850: synonym of Bostryx Troschel, 1847
• Bilamelliferus Weyrauch, 1958: synonym of Bostryx Troschel, 1847 (junior synonymy)
• Cochlogena Férussac, 1821: synonym of Bulimulus Leach, 1814 (junior synonym)
• Diaphanomormus Weyrauch, 1964: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
• Elatibostryx Weyrauch, 1958: synonym of Bostryx Troschel, 1847
• Floreziellus Weyrauch, 1967: synonym of Bostryx Troschel, 1847 (junior synonymy)
• Geoceras Pilsbry, 1896: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
• Globulinus Crosse & P. Fischer, 1875: synonym of Rabdotus Albers, 1850
• Goniognathmus Crosse & P. Fischer, 1875: synonym of Drymaeus Albers, 1850 (junior synonym)
• Hamadryas Albers, 1850: synonym of Drymaeus Albers, 1850
• Hannarabdotus Emerson & Jacobson, 1964: synonym of Rabdotus Albers, 1850
• Kionoptyx Haas, 1966: synonym of Bostryx Troschel, 1847 (junior synonymy)
• Kuschelenia Hylton Scott, 1951: synonym of Bocourtia (Kuschelenia) Hylton Scott, 1951 represented as Bocourtia Rochebrune, 1882
• Leptodrymaeus Pilsbry, 1946: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
• Leptomerus Albers, 1850: synonym of Bulimulus Leach, 1814
• Leptomormus Weyrauch, 1958: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
• Liostracus [sic]: synonym of Leiostracus Albers, 1850 (misspelling of original genus, Leiostracus Albers, 1850)
• Loboa Ihering, 1917: synonym of Bulimulus Leach, 1814
• Mormus E. von Martens, 1860: synonym of Drymaeus Albers, 1850
• Multifasciatus Weyrauch, 1958: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
• Naesiotellus Weyrauch, 1967: synonym of Bostryx Troschel, 1847 (junior synonym)
• Navicula Spix, 1827: synonym of Cochlorina Jan, 1830
• Nesiotes E. von Martens, 1860: synonym of Naesiotus Albers, 1850
• Obstrussus Parodiz, 1946: synonym of Naesiotus Albers, 1850
• Olinodia Dall, 1920: synonym of Naesiotus (Reclasta) Dall, 1920 represented as Naesiotus Albers, 1850
• Orphaicus Schaufuss, 1869: synonym of Thaumastus E. von Martens, 1860
• Orphnus Albers, 1850: synonym of Thaumastus E. von Martens, 1860
• Orthotomium Crosse & P. Fischer, 1875: synonym of Rabdotus Albers, 1850
• Pachyotus H. Beck, 1837: synonym of Auris Spix, 1827
• Pachytholus Strebel, 1909: synonym of Thaumastus E. von Martens, 1860
• Pampasinus Weyrauch, 1958: synonym of Bostryx (Platybostryx) Pilsbry, 1896: synonym ofBostryx Troschel, 1847
• Peronaeus Albers, 1850: synonym of Bostryx Troschel, 1847
• Phenacotaxus Dall, 1912: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
• Plecochilus Agassiz, 1846: synonym of Plekocheilus Guilding, 1827 (unjustified emendation)
• Pseudoperonaeus Weyrauch, 1958: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
• Pseudorhodea Dall, 1895: synonym of Rabdotus (Plicolumna) J.G. Cooper, 1895 represented as Rabdotus Albers, 1850
• Pyrgus Albers, 1850: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
• Rhabdotus [sic]: synonym of Rabdotus Albers, 1850 (incorrect subsequent spelling)
• Rhaphiellus L. Pfeiffer, 1856: synonym of Naesiotus Albers, 1850
• Rimatula Parodiz, 1946: synonym of Protoglyptus Pilsbry, 1897
• Saeronia Dall, 1920: synonym of Naesiotus Albers, 1850
• Semiclausaria L. Pfeiffer, 1856: synonym of Drymaeus Albers, 1850 (junior synonym)
• Siphalomphix Rafinesque, 1833: synonym of Bulimulus Leach, 1814
• Sonorina Pilsbry, 1896: synonym of Rabdotus (Leptobyrsus) Crosse & P. Fischer, 1875 represented as Rabdotus Albers, 1850
• Spiroscutalus Pilsbry, 1932: synonym of Scutalus Albers, 1850
• Stenostoma Spix, 1827: synonym of Anctus E. von Martens, 1860
• Tatutor Jousseaume, 1887: synonym of Thaumastus E. von Martens, 1860
• Teneritia J. Mabille, 1898: synonym of Spartocentrum Dall, 1895
• Tholus Strebel, 1909: synonym of Thaumastus E. von Martens, 1860
• Xenothauma Fulton 1896:^[17][18] synonym of Scutalus Albers, 1850
• Zaplagius Pilsbry, 1896: synonym of Cochlorina Jan, 1830

This article incorporates CC-BY-3.0 text from the reference^[7]

• Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

Wikimedia Commons has media related to Bulimulidae.

• Neubert, E.; Janssen, R. (2004). "Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae". Archiv für Molluskenkunde. 133: 193–297. doi:10.1127/arch.moll/133/2004/193.
• Breure, Abraham S.H.; Ablett, Jonathan D. (21 March 2014). "Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London". ZooKeys (392): 1–367. Bibcode:2014ZooK..392....1B. doi:10.3897/zookeys.392.6328. PMC 3974435. PMID 24715782.