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Charruodon Temporal range: Carnian ~235–222 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Synapsida
Clade:	Therapsida
Clade:	Cynodontia
Clade:	Probainognathia
Genus:	†Charruodon Abdala & Ribeiro, 2000
Species:	†C. tetracuspidatus
Binomial name
†Charruodon tetracuspidatus Abdala & Ribeiro, 2000

Charruodon is an extinct genus of cynodonts which existed in the Hyperodapedon Assemblage Zone of the Santa Maria Formation in the Paraná Basin in southeastern Brazil during the Late Triassic. The genus contains only the type species Charruodon tetracuspidatus, which is known from a single specimen of uncertain provenance.^[1]^[2] Upon its first description, Charruodon was tentatively placed within the family Therioherpetidae, but a 2017 study by Agustín G. Martinelli and colleagues instead recovered it as a more basal member of Probainognathia.^[3] In 2023, Hoffmann, Ribeiro & de Andrade reinterpreted the specimen as representing an early ontogenetic stage, and C. tetracuspidatus to be a nomen dubium.^[4]

References

^ Abdala, F.; Ribeiro, A. M. (2000). "A new therioherpetid cynodont from the Santa Maria Formation (middle Late Triassic), southern Brazil" (PDF). Geodiversitas. 22 (4): 589–596. Archived from the original (PDF) on 2011-06-12. Retrieved 2010-04-05.
^ Langer, M. C. (2005). "Studies on continental Late Triassic tetrapod biochronology. I. The type locality of Saturnalia tupiniquim and the faunal succession in south Brazil" (PDF). Journal of South American Earth Sciences. 19: 205–218. doi:10.1016/j.jsames.2005.04.003. Archived from the original (PDF) on 2010-09-24.
^ Martinelli, A. G.; Eltink, E.; Da-Rosa, Á. A. S.; Langer, M. C. (2017). "A new cynodont from the Santa Maria formation, south Brazil, improves Late Triassic probainognathian diversity". Papers in Palaeontology. 3 (3): 401–423. doi:10.1002/spp2.1081.
^ Hoffmann, C. A.; Ribeiro, A. M.; de Andrade, M. B. (2023). "On the dentition, tooth replacement, and taxonomic status of Charruodon tetracuspidatus Abdala & Ribeiro, 2000: A bizarre cynodont from the middle upper Triassic of southern Brazil". The Anatomical Record. doi:10.1002/ar.25349. PMID 37950602.

Cynodontia

Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Eutheriodontia

Synapsida

see Synapsida
Cynodontia	see below↓

Cynodontia

†Dvinia †Nanocynodon †Madysaurus †Uralocynodon
†Charassognathidae	Abdalodon Charassognathus Nshimbodon
†Procynosuchidae	Procynosuchus Sludica

Epicynodontia

†Galesauridae

Galesaurus
Progalesaurus

Eucynodontia

†Kataigidodon
†Cynognathia	see Cynognathia
Probainognathia	see below↓

Probainognathia

†Agudotherium †Candelariodon †Charruodon †Lumkuia †Protheriodon †Tessellatia
†Chiniquodontidae	Aleodon Chiniquodon Cromptodon? Riojanodon
†Probainognathidae	Bonacynodon Probainognathus
†Ecteniniidae	Diegocanis Ecteninion Trucidocynodon

Prozostrodontia

†Prozostrodontidae

Prozostrodon
Pseudotherium

†Tritheledontidae

Mammaliamorpha

†Adelobasileus †Botucaraitherium †Brasilodon †Sinoconodon †Tikiodon
†Tritylodontidae	Bienotherium Bienotheroides Bocatherium Dianzhongia Dinnebitodon Fossiomanus Kayentatherium Lufengia Montirictus Nuurtherium Oligokyphus Polistodon Shartegodon Stereognathus Tritylodon Xenocretosuchus Yuanotherium Yunnanodon
Mammaliaformes	see below↓

Mammaliaformes

†Kuehneotheriidae

†Shuotheriidae

†Docodonta

†Haramiyida

Euharamiyida

Allostaffia? Arboroharamiya Baidabatyr? Butlerodon Cryoharamiya Kermackodon? Maiopatagium Megaconus? Qishou Sharypovoia Shenshou Sineleutherus Vilevolodon Woodeatonia Xianshou Gondwanatheria?
Hahnodontidae?	Cifelliodon Denisodon Hahnodon

Crown-Mammalia

- see Mammalia