Urocyon

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Urocyon

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(Redirected from Cozumel fox)

Urocyon^[2] Temporal range: 10.3–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N ^[1]

Gray and island fox
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Canidae
Subfamily:	Caninae
Genus:	Urocyon Baird, 1857^[3]
Type species
Canis virginianus^[5] Schreber, 1774^[6]
Species
U. cinereoargenteus (Schreber), 1774 U. littoralis Baird, 1857 † U. citrinus Tedford, Wang & Taylor (2009)^[4] † U. galushai Tedford, Wang & Taylor (2009)^[4] † U. minicephalus Martin, 1974 † U. progressus Stevens, 1965 † U. webbi Tedford, Wang & Taylor (2009)^[4]

Urocyon (Greek: "tailed dog"^[7]) is a genus of Canidae which includes the gray fox (Urocyon cinereoargenteus) and the island fox (Urocyon littoralis).^[2] These two fox species are found in the Western Hemisphere. Whole genome sequencing indicates that Urocyon is the most basal genus of the living canids.^[8]^{[failed verification]} Fossils of what is believed to be the ancestor of the gray fox, Urocyon progressus, have been found in Kansas and date to the Upper Pliocene,^[9] with some undescribed specimens dating even older.^[10]

Extant species

[edit]

Genus **Urocyon** – Baird, 1857 – two species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Gray fox	Urocyon cinereoargenteus (Schreber, 1775)	Southern half of North America from southern Canada to the northern part of South America (Venezuela and Colombia), excluding the mountains of northwestern United States	Size: Habitat: Diet:	LC
Island fox	Urocyon littoralis (Baird, 1857) Six subspecies U. l. littoralis (the nominate subspecies) of San Miguel Island, U. l. dickeyi of San Nicolas Island, U. l. catalinae of Santa Catalina Island, U. l. clementae of San Clemente Island, U. l. santacruzae of Santa Cruz Island, and U. l. santarosae of Santa Rosa Island.	Channel Islands (off the coast of Southern California)	Size: Habitat: Diet:	NT

Extinct species

[edit]

Species name	Type specimen era and location^[4]
U. citrinus	Tedford et al., 2009. Early Irvingtonian, Citrus County, Florida.
U. galushai	Tedford et al., 2009. Late Blancan, San Simon Valley, Graham County, Arizona.
U. minicephalus	Martin, 1974. Late Irvingtonian, Sumter County, Florida.
U. progressus	Stevens, 1965. Early Blancan, Meade County, Kansas. A later review found that the material represents Urocyon, but because of its fragmentary nature cannot be diagnosed as to species.^[4]
U. webbi	Tedford et al., 2009. Middle Hemphillian, Citrus County, Florida.

Cozumel fox

[edit]

The Cozumel fox is a critically endangered or extinct small gray fox found on the island of Cozumel, Mexico. The last reported sighting was in 2001, but surveys focusing on this species have not yet been carried out.^[11]

The Cozumel fox has not been scientifically described, but is a dwarf form as is the island fox, but slightly larger being up to three-quarters the size of the gray fox. No skins or complete skulls of the Cozumel fox exist in any museum exhibitions, so scientists have mainly examined sub-fossils collected during archaeological excavations of Mayan civilizations who inhabited the island about 1,500–500 years ago. Upon evaluating bones from about 12 adult individuals, scientists have concluded that the Cozumel fox is extremely small – approximately 60-80% the body size of other mainland specimens.^[12]

The fox had been isolated on the island of Cozumel for at least 5,000 years, and probably far longer. This would indicate that Urocyon had colonized the island before the first arrival of humans there.^[12]

References

[edit]

^ "Fossilworks: Urocyon".
^ ^a ^b Wozencraft, W. C. (2005). "Order Carnivora". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 582–583. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ Baird, S. 1857. Reports of explorations and surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean. vol.8(1):121, 138. Vulpinae
^ ^a ^b ^c ^d ^e Tedford, Richard H.; Wang, Xiaoming; Taylor, Beryl E. (2009). "Phylogenetic systematics of the North American fossil Caninae (Carnivora: Canidae)" (PDF). Bulletin of the American Museum of Natural History. 325: 1–218. doi:10.1206/574.1. hdl:2246/5999. S2CID 83594819.
^ Wilson, D. E.; Reeder, D. M., eds. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ Schreber, Johann Christian Daniel; Goldfuss, Georg August; Wagner, Andreas Johann (1774). Die Säugthiere in Abbildungen nach der Natur, mit Beschreibungen. Vol. 3. p. 361. doi:10.5962/bhl.title.67399.( The mammals in illustrations according to nature, with descriptions)
^ Urocyon.
^ Lindblad-Toh, K.; Wade, C.M.; Mikkelsen, T.S.; Karlsson, E.K.; Jaffe, D.B.; Kamal, M.; et al. (2005). "Genome sequence, comparative analysis and haplotype structure of the domestic dog". Nature. 438 (7069): 803–819. Bibcode:2005Natur.438..803L. doi:10.1038/nature04338. PMID 16341006.
^ Stevens, M.S. (1965). "A new species of Urocyon from the Upper Pliocene of Kansas". Journal of Mammalogy. 46 (2): 265–269. doi:10.2307/1377846. JSTOR 1377846.
^ Prevosti, F.J.; Rincóon, A.D. (2007). "A new fossil canid assemblage from the late Pleistocene of northern South America: The canids of the Inciarte asphalt pit (Zulia, Venezuela), fossil record and biogeography". J. Paleontol. 81 (5): 1053–1065. doi:10.1666/pleo05-143.1. hdl:11336/242701. S2CID 131259363.
^ Cuarón, Alfredo D.; Martinez-Morales, Miguel Angel; McFadden, Katherine W.; Valenzuela, David & Gompper, Matthew E. (2004). "The status of dwarf carnivores on Cozumel Island, Mexico". Biodiversity and Conservation. 13 (2): 317–331. CiteSeerX 10.1.1.511.2040. doi:10.1023/B:BIOC.0000006501.80472.cc. S2CID 25730672.
^ ^a ^b Gompper, M.E.; Petrites, A.E.; Lyman, R.L. (2006). "Cozumel Island fox (Urocyon sp.) dwarfism and possible divergence history based on subfossil bones". Journal of Zoology. 270 (1): 72–77. doi:10.1111/j.1469-7998.2006.00119.x. ISSN 1469-7998.

Extant Carnivora species

Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Eutheria
Superorder: Laurasiatheria

Feliformia ("cat-like" carnivorans)

Feliformia

African palm civet (Nandinia binotata)

Feloidea

Prionodon (Asiatic linsangs)

Felidae (cats)

Pantherinae

Neofelis	Sunda clouded leopard (N. diardi) Clouded leopard (N. nebulosa)
Panthera	Snow leopard (P. uncia) Tiger (P. tigris) Jaguar (P. onca) Lion (P. leo) Leopard (P. pardus)

Felinae sensu stricto

Bay cat
lineage

Pardofelis	Marbled cat (P. marmorata)
Catopuma	Bay cat (C. badia) Asian golden cat (C. temminckii)

Caracal
lineage

Serval (Leptailurus serval)
Caracal	African golden cat (C. aurata) Caracal (C. caracal)

Leopardus

Lynx

Puma
lineage

Acinonyx	Cheetah (A. jubatus)
Jaguarundi (Herpailurus yagouaroundi)
Puma	Cougar (P. concolor)

Leopard cat
lineage

Pallas's cat (Otocolobus manul)
Prionailurus	Leopard cat (P. bengalensis) Sunda leopard cat (P. javanensis) Flat-headed cat (P. planiceps) Rusty-spotted cat (P. rubiginosus) Fishing cat (P. viverrinus)

Felis

Viverroidea

- see below↓

Binturong (Arctictis binturong) Small-toothed palm civet (Arctogalidia trivirgata) Sulawesi palm civet (Macrogalidia musschenbroekii) Masked palm civet (Paguma larvata)
Paradoxurus	Asian palm civet (P. hermaphroditus) Brown palm civet (P. jerdoni) Golden palm civet (P. zeylonensis)

Viverrinae
sensu lato

Viverrinae
sensu stricto

Small Indian civet (Viverricula indica) African civet (Civettictis civetta)
Viverra	Malabar large-spotted civet (V. civettina) Large-spotted civet (V. megaspila) Malayan civet (V. tangalunga) Large Indian civet (V. zibetha)

Genettinae

Poiana
(African linsangs)

Genetta
(genets)

Herpestoidea	see below↓

Herpestoidea

Hyaenidae
(hyenas)

Proteles

Aardwolf (P. cristata)

Hyaeninae
(bone-crushing hyenas)

Striped hyena (Hyaena hyaena) Brown hyena (Parahyaena brunnea)
Crocuta	Spotted hyena (C. crocuta)

Herpestidae sensu lato

Eupleridae
(Malagasy
carnivorans)

Euplerinae

Fossa (Cryptoprocta ferox) Malagasy civet (Fossa fossana)
Eupleres	Eastern falanouc (E. goudotii) Western falanouc (E. major)

Galidiinae

Ring-tailed vontsira (Galidia elegans)
Galidictis	Broad-striped Malagasy mongoose (G. fasciata) Grandidier's mongoose (G. grandidieri)
Narrow-striped mongoose (Mungotictis decemlineata)
Salanoia	Brown-tailed mongoose (S. concolor) Durrell's vontsira (S. durrelli)

Herpestidae
sensu stricto
(mongooses)

Mungotinae

Suricata	Meerkat (S. suricatta)
Liberian mongoose (Liberiictus kuhni)
Mungos	Gambian mongoose (M. gambianus) Banded mongoose (M. mungo)
Pousargues's mongoose (Dologale dybowskii)
Helogale	Ethiopian dwarf mongoose (H. hirtula) Common dwarf mongoose (H. parvula)
Crossarchus	Alexander's kusimanse (C. alexandri) Angolan kusimanse (C. ansorgei) Common kusimanse (C. obscurus) Flat-headed kusimanse (C. platycephalus)

Herpestinae

Marsh mongoose (Atilax paludinosus) Long-nosed mongoose (Xenogale naso)
Urva	Small Indian mongoose (U. auropunctata) Short-tailed mongoose (U. brachyura) Indian grey mongoose (U. edwardsii) Indian brown mongoose (U. fusca) Javan mongoose (U. javanica) Collared mongoose (U. semitorquata) Ruddy mongoose (U. smithii) Crab-eating mongoose (U. urva) Stripe-necked mongoose (U. vitticolla)
White-tailed mongoose (Ichneumia albicauda) Yellow mongoose (Cynictis penicillata) Selous's mongoose (Paracynictis selousi) Meller's mongoose (Rhynchogale melleri)
Bdeogale	Bushy-tailed mongoose (B. crassicauda) Jackson's mongoose (B. jacksoni) Black-footed mongoose (B. nigripes)
Herpestes	Angolan slender mongoose (H. flavescens) Egyptian mongoose (H. ichneumon) Somalian slender mongoose (H. ochracea) Cape gray mongoose (H. pulverulenta) Common slender mongoose (H. sanguinea)

Caniformia ("dog-like" carnivorans)

Canidae (dogs)

Urocyon

Vulpini

Bat-eared fox (Otocyon megalotis)
Nyctereutes	Common raccoon dog (N. procyonoides) Japanese raccoon dog (N. viverrinus)
Vulpes (foxes)	Bengal fox (V. bengalensis) Blanford's fox (V. cana) Cape fox (V. chama) Corsac fox (V. corsac) Tibetan fox (V. ferrilata) Arctic fox (V. lagopus) Kit fox (V. macrotis) Pale fox (V. pallida) Rüppell's fox (V. rueppelli) Swift fox (V. velox) Red fox (V. vulpes) Fennec fox (V. zerda)

Canini
(true dogs)

Cerdocyonina
(zorro)

Maned wolf (Chrysocyon brachyurus)
Speothos	Bush dog (S. venaticus)
Short-eared dog (Atelocynus microtis) Crab-eating fox (Cerdocyon thous)
Lycalopex	Culpeo (L. culpaeus) Darwin's fox (L. fulvipes) South American gray fox (L. griseus) Pampas fox (L. gymnocercus) Sechuran fox (L. sechurae) Hoary fox (L. vetulus)

Canina
(wolf-like canids)

Lupulella	Side-striped jackal (L. adustus) Black-backed jackal (L. mesomelas)
African wild dog (Lycaon pictus) Dhole (Cuon alpinus)
Canis	Golden jackal (C. aureus) Domestic dog (C. familiaris) Coyote (C. latrans) African wolf (C. lupaster) Wolf (C. lupus) Eastern wolf (C. lycaon) Red wolf (C. rufus) Ethiopian wolf (C. simensis)

Arctoidea

Ursidae
(bears)

Ailuropoda

Giant panda (A. melanoleuca)

Tremarctos

Spectacled bear (T. ornatus)

Ursinae

Sloth bear (Melursus ursinus) Sun bear (Helarctos malayanus)
Ursus	American black bear (U. americanus) Asian black bear (U. thibetanus) Brown bear (U. arctos) Polar bear (U. maritimus)

Mustelida

Pinnipedia (seals)	see below↓
Musteloidea	see below↓

Pinnipedia (seals)

Odobenidae

Walrus (Odobenus rosmarus)

Otariidae
(eared seals:
fur seals,
sea lions)

Callorhinus

Northern fur seal (C. ursinus)

Otariinae

Steller sea lion (Eumetopias jubatus)
Zalophus	California sea lion (Z. californianus) Galápagos sea lion (Z. wollebaeki)

South American sea lion (Otaria flavescens)

Neophoca

Australian sea lion (N. cinerea)

New Zealand sea lion (Phocarctos hookeri)

Arctocephalus

Phocidae
(earless seals
or true seals)

Phocinae
("northern seals")

Phocini

Harp seal (Pagophilus groenlandicus) Ribbon seal (Histriophoca fasciata) Grey seal (Halichoerus grypus)
Phoca	Spotted seal (P. largha) Harbor seal (P. vitulina)
Pusa	Caspian seal (P. caspica) Ringed seal (P. hispida) Baikal seal (P. sibirica)

Monachinae
("southern seals")

Monachini
(monk seals)

Mediterranean monk seal (Monachus monachus)
Neomonachus	Hawaiian monk seal (N. schauinslandi)

Mirounga
(elephant seals)

Lobodontini
(Antarctic seals)

Musteloidea

Ailuridae

Red panda (Ailurus fulgens)

Mephitidae
(skunks)

Conepatus (hog-nosed skunks)	Molina's hog-nosed skunk (C. chinga) Humboldt's hog-nosed skunk (C. humboldtii) American hog-nosed skunk (C. leuconotus) Striped hog-nosed skunk (C. semistriatus)
Mephitis	Hooded skunk (M. macroura) Striped skunk (M. mephitis)
Mydaus (stink badgers)	Sunda stink badger (M. javanensis) Palawan stink badger (M. marchei)
Spilogale (spotted skunks)	Southern spotted skunk (S. angustifrons) Western spotted skunk (S. gracilis) Eastern spotted skunk (S. putorius) Pygmy spotted skunk (S. pygmaea)

Procyonidae

Kinkajou (Potos flavus)

Bassariscus

Procyon
(raccoons)

Bassaricyon
(olingos)

Nasuina
(coatis)

Nasua	South American coati (N. nasua) White-nosed coati (N. narica)
Nasuella	Eastern mountain coati (N. meridensis) Western mountain coati (N. olivacea)

Mustelidae

- see below↓

Mustelidae

American badger (Taxidea taxus)

Mellivora

Honey badger (M. capensis)

Melinae
(Eurasian badgers)

Arctonyx	Northern hog badger (A. albogularis) Greater hog badger (A. collaris) Sumatran hog badger (A. hoevenii)
Meles	Japanese badger (M. anakuma) Caucasian badger (M. canescens) Asian badger (M. leucurus) European badger (M. meles)

Melogale
(ferret-badgers)

Guloninae

Tayra (Eira barbara)
Pekania	Fisher (P. pennanti)
Gulo	Wolverine (G. gulo)
Martes (martens)	American marten (M. americana) Pacific marten (M. caurina) Yellow-throated marten (M. flavigula) Beech marten (M. foina) Nilgiri marten (M. gwatkinsii) European pine marten (M. martes) Japanese marten (M. melampus) Sable (M. zibellina)

Ictonychinae

Lyncodontini

Patagonian weasel (Lyncodon patagonicus)
Galictis (grisons)	Lesser grison (G. cuja) Greater grison (G. vittata)

Ictonychini
(African polecats)

Vormela	Marbled polecat (V. peregusna)
African striped weasel (Poecilogale albinucha)
Ictonyx	Saharan striped polecat (I. libycus) Striped polecat (I. striatus)

Lutrinae
(otters)

Giant otter (Pteronura brasiliensis)
Lontra	North American river otter (L. canadensis) Marine otter (L. felina) Neotropical otter (L. longicaudis) Southern river otter (L. provocax)
Enhydra	Sea otter (E. lutris)
Spotted-necked otter (Hydrictis maculicollis)
Lutra	Eurasian otter (L. lutra) Hairy-nosed otter (L. sumatrana)
Lutrogale	Smooth-coated otter (L. perspicillata)
Aonyx	African clawless otter (A. capensis) Asian small-clawed otter (A. cinereus) Congo clawless otter (A. congicus)

Mustelinae

Neogale

Mustela
(weasels)

Taxon identifiers
Urocyon	Wikidata: Q738191 Wikispecies: Urocyon ADW: Urocyon BOLD: 3080 CoL: 8583 EoL: 20188 EPPO: 1URCYG GBIF: 2434558 iNaturalist: 42075 IRMNG: 1313570 ITIS: 180608 MSW: 14000840 NCBI: 55039 Open Tree of Life: 264179 Paleobiology Database: 41247

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