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Protathlitis - Wikipedia

From Wikipedia, the free encyclopedia

Protathlitis Temporal range: Early Cretaceous, Barremian 129.4–125 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓

Skeletal diagram of the related Baryonyx, showing the known bones of Protathlitis in red
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Clade:	Theropoda
Family:	†Spinosauridae
Subfamily:	†Baryonychinae
Genus:	†Protathlitis Santos-Cubedo et al., 2023
Type species
†Protathlitis cinctorrensis Santos-Cubedo et al., 2023

Protathlitis (meaning "champion") is a genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Arcillas de Morella Formation of Castellón, Spain. The type and only species is Protathlitis cinctorrensis, known from a partial skeleton. It was a basal member of the Baryonychinae.^[1] Its discovery, as well as those of the spinosaurids Camarillasaurus, Iberospinus, Riojavenatrix, and the contemporary Vallibonavenatrix shows that the Iberian Peninsula held a diverse assemblage of spinosaurids during the Early Cretaceous.^[2]

Discovery and naming

The holotype remains, the maxillary fragment 8ANA-109 and caudal vertebrae 3ANA83, 4ANA43, 4ANA69, 4ANA76, and 5ANA78, were recovered from the ANA site of the Arcillas de Morella Formation, which was discovered in 1998 and remained unexplored until 2002. A tooth, 4ANA-11, possibly from the left mandible or right maxilla, was also referred.

The remains were described as a new genus and species of spinosaurid in 2023, Protathlitis cinctorrensis. The genus name, "Protathlitis", is Greek for "champion", and is dedicated to the 2020–21 UEFA Europa League won by Villareal C.F. and in light of the club's centenary in 2023. The specific name, "cinctorrensis", honors Cinctorres, the town where the remains were discovered.^[1]

In 2024, Montealegre, Castillo-Visa & Sellés tentatively assigned specimen IPS919, a nearly complete tooth including a partial root from the Arcillas de Morella Formation, to cf. Protathlitis.^[3]

Classification

Santos-Cubedo et al. (2023) performed a phylogenetic analysis, placing Protathlitis as the basalmost member the Baryonychinae. Their results are shown in the cladogram below:^[1]

Spinosauridae

Camarillasaurus

Baryonychinae

Protathlitis

Baryonyx

	Ceratosuchops

	Suchomimus

Spinosaurinae

Ichthyovenator

Vallibonavenatrix

	Spinosaurus

	Irritator

Palaeoenvironment

Protathlitis hails from the Arcillas de Morella Formation, which has been dated to the Barremian stage of the Early Cretaceous period, between 129.4 and 125 million years ago. It coexisted in this environment with other dinosaurs including the ornithischians Iguanodon bernissartensis and Morelladon beltrani, an indeterminate sauropod, and the fellow spinosaurid Vallibonavenatrix.^[4]^[5]^[6]

References

^ ^a ^b ^c Santos-Cubedo, A.; de Santisteban, C.; Poza, B.; Meseguer, S. (2023). "A new spinosaurid dinosaur species from the Early Cretaceous of Cinctorres (Spain)". Scientific Reports. 13 (1). 6471. doi:10.1038/s41598-023-33418-2. hdl:10234/203142.
^ Isasmendi, E.; Cuesta, E.; Díaz-Martínez, I.; Company, J.; Sáez-Benito, P.; Viera, L. I.; Torices, A.; Pereda-Suberbiola, P. (2024). "Increasing the theropod record of Europe: a new basal spinosaurid from the Enciso Group of the Cameros Basin (La Rioja, Spain). Evolutionary implications and palaeobiodiversity". Zoological Journal of the Linnean Society. doi:10.1093/zoolinnean/zlad193. {{cite journal}}: |last4= has generic name (help)
^ Montealegre, Adrian; Castillo-Visa, Oscar; Sellés, Albert (2024-01-31). "New theropod remains from the late Barremian (Early Cretaceous) of Eastern Iberian Peninsula". Historical Biology: 1–11. doi:10.1080/08912963.2024.2308220. ISSN 0891-2963.
^ Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Cretaceous, Europe)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 562. ISBN 0-520-24209-2.
^ Verdú, F.J.; Godefroit, P.; Royo-Torres, R.; Cobos, A.; Alcalá, L. (2017). "Individual variation in the postcranial skeleton of the Early Cretaceous Iguanodon bernissartensis (Dinosauria: Ornithopoda)". Cretaceous Research. 74: 65–86. doi:10.1016/j.cretres.2017.02.006.
^ Malafaia, E.; Miguel Gasulla, J.; Escaso, F.; Narváez, I.; Luis Sanz, J.; Ortega, F. (2019). "A new spinosaurid theropod (Dinosauria: Megalosauroidea) from the late Barremian of Vallibona, Spain: Implications for spinosaurid diversity in the Early Cretaceous of the Iberian Peninsula". Cretaceous Research. 106: 104221. doi:10.1016/j.cretres.2019.104221. S2CID 202189246.

Theropoda

Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Clade: Avemetatarsalia
Clade: Dinosauria

Avemetatarsalia

see Avemetatarsalia
Theropoda	see below↓

Theropoda

Neotheropoda

Coelophysoidea

Dracoraptor? Panguraptor Pendraig Podokesaurus Powellvenator
Coelophysidae	Camposaurus Coelophysis Lucianovenator Megapnosaurus? Panguraptor? Procompsognathus Pterospondylus? Segisaurus

Averostra

- see below↓

Dubious neotheropods

Teinurosaurus
Halticosaurus

Averostra

Dornraptor
Lophostropheus?

Ceratosauria

Berberosaurus Fosterovenator? Jubbulpuria Lukousaurus? Saltriovenator
Ceratosauridae	Ceratosaurus Genyodectes

Abelisauroidea

Noasauridae

Afromimus? Austrocheirus? Berthasaura Compsosuchus? Dahalokely? Deltadromeus? Genusaurus? Kiyacursor Ligabueino Spinostropheus
Elaphrosaurinae	Elaphrosaurus Huinculsaurus Limusaurus
Noasaurinae	Laevisuchus Masiakasaurus Noasaurus Velocisaurus Vespersaurus

Abelisauridae

Majungasaurinae

Carnotaurinae

Majungasaurinae?

Brachyrostra

Dahalokely? Ekrixinatosaurus Elemgasem Guemesia Ilokelesia Llukalkan Niebla Rahiolisaurus? Skorpiovenator Thanos
Furileusauria	Abelisaurus Aucasaurus Caletodraco Carnotaurus Elemgasem? Koleken Pycnonemosaurus Quilmesaurus Viavenator

Tetanurae	see below↓

Tetanurae

Orionides

Megalosauroidea

Yunyangosaurus
Piatnitzkysauridae	Condorraptor Marshosaurus Piatnitzkysaurus Xuanhanosaurus?

Megalosauria

Streptospondylus?

Megalosauridae

Eustreptospondylus
Megalosaurinae	Duriavenator Megalosaurus Torvosaurus Wiehenvenator
Afrovenatorinae	Afrovenator Dubreuillosaurus Leshansaurus Magnosaurus Piveteausaurus Poekilopleuron?

Spinosauridae

Baryonychinae

Baryonyx Protathlitis Riojavenatrix Suchosaurus? Vallibonavenatrix?
Ceratosuchopsini	Ceratosuchops Riparovenator Suchomimus

Spinosaurinae

Camarillasaurus? Ichthyovenator Irritator Oxalaia Siamosaurus
Spinosaurini	Sigilmassasaurus Spinosaurus

Avetheropoda	see below↓

Avetheropoda

Gasosaurus?
Lourinhanosaurus?

Carnosauria

Allosauroidea

Metriacanthosauridae

Xuanhanosaurus? Yangchuanosaurus
Metriacanthosaurinae	Alpkarakush Metriacanthosaurus Shidaisaurus Siamotyrannus Sinraptor

Carcharodontosauridae

Carcharodontosaurinae

Carcharodontosaurus
Giganotosaurini	Giganotosaurus Mapusaurus Meraxes Taurovenator Tyrannotitan

Megaraptora?

Aoniraptor Bahariasaurus? Deltadromeus? Fukuiraptor Phuwiangvenator Rapator Siats? Chilantaisaurus?
Megaraptoridae	Aerosteon Australovenator Maip Megaraptor Murusraptor Orkoraptor Tratayenia

Coelurosauria

- see below↓

Coelurosauria

Tyrannoraptora

Tyrannosauroidea

Chingkankousaurus Labocania?
Coeluridae?	Coelurus Tanycolagreus?
Proceratosauridae	Guanlong Kileskus Proceratosaurus Sinotyrannus Yutyrannus

Albertosaurus
Gorgosaurus

Tyrannosaurinae

Asiatyrannus Nanuqsaurus
Alioramini	Alioramus Qianzhousaurus
Daspletosaurini	Daspletosaurus Thanatotheristes
Teratophoneini	Bistahieversor? Dynamoterror Labocania? Lythronax Teratophoneus
Tyrannosaurini	Tarbosaurus Tyrannosaurus Zhuchengtyrannus

Maniraptoromorpha	see below↓

Dubious coelurosaurs	Iliosuchus Kakuru Phaedrolosaurus Shanyangosaurus

Maniraptoromorpha

Neocoelurosauria

Aniksosaurus
Compsognathidae	Aniksosaurus? Aristosuchus Beipiaognathus? Compsognathus Huaxiagnathus Mirischia Scipionyx? Sinocalliopteryx Sinosauropteryx Xunmenglong

Maniraptoriformes

Compsognathidae?

Ornithomimosauria

Macrocheiriformes

Harpymimus Pelecanimimus Shenzhousaurus
Deinocheiridae	Beishanlong Deinocheirus Garudimimus Harpymimus? Paraxenisaurus? Tyrannomimus
Ornithomimidae	Aepyornithomimus Anserimimus Archaeornithomimus "Coelosaurus" Dromiceiomimus Gallimimus Ornithomimus Qiupalong Rativates Sinornithomimus Struthiomimus Tototlmimus

Maniraptora

- see Maniraptora
- includes birds

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