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Theriimorpha Temporal range: Toarcian–Present PreꞒ Ꞓ O S D C P T J K Pg N

Skeleton of Gobiconodon ostromi (Eutriconodonta)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Clade:	Theriimorpha Rowe, 1993
Subgroups
†Eutriconodonta †Fruitafossor? Theriiformes Trechnotheria

Theriimorpha is a clade of mammals defined as including all mammals more closely related to therians (including placentals and marsupials) than to monotremes. Eutriconodonta is usually considered among the most basal members of this group, with other members more closely related to therians like Allotherians placed in the subclade Theriiformes,^[1] though Eutriconodonta has also been recovered as less closely related to therians than monotremes are in some analyses, placing them outside the crown group of Mammalia.^[2]^[3] The unusual Late Jurassic digging mammal Fruitafossor has also been suggested to be a basal theriimorph in some studies.^[4] The position of Allotheria (which contains the multituberculates, among others) within Theriimorpha is controversial. While many studies recover Allotheria within crown Mammalia as more closely related to therians than to monotreme as a member of the clade Theriiformes, some studies recover the group as outside of crown Mammalia.^[5]

References

^ Rowe T (1993). "Phylogenetic Systematics and the Early History of Mammals". In Szalay FS, Novacek MJ, McKenna MC (eds.). Mammal Phylogeny. New York, NY: Springer New York. pp. 129–145. doi:10.1007/978-1-4613-9249-1_10. ISBN 978-1-4613-9251-4.
^ Luo, Zhe-Xi; Gatesy, Stephen M.; Jenkins, Farish A.; Amaral, William W.; Shubin, Neil H. (2015-12-22). "Mandibular and dental characteristics of Late Triassic mammaliaform Haramiyavia and their ramifications for basal mammal evolution". Proceedings of the National Academy of Sciences. 112 (51): E7101-9. Bibcode:2015PNAS..112E7101L. doi:10.1073/pnas.1519387112. ISSN 0027-8424. PMC 4697399. PMID 26630008.
^ Hoffmann, Simone; Beck, Robin M. D.; Wible, John R.; Rougier, Guillermo W.; Krause, David W. (2020-12-14). "Phylogenetic placement of Adalatherium hui (Mammalia, Gondwanatheria) from the Late Cretaceous of Madagascar: implications for allotherian relationships". Journal of Vertebrate Paleontology. 40 (sup1): 213–234. Bibcode:2020JVPal..40S.213H. doi:10.1080/02724634.2020.1801706. ISSN 0272-4634. S2CID 230968231.
^ Hughes EM, Wible JR, Spaulding M, Luo ZX (May 2015). "Mammalian petrosal from the Upper Jurassic Morrison Formation of Fruita, Colorado". Annals of Carnegie Museum. 83 (1): 1–17. doi:10.2992/007.083.0101. S2CID 83598504.
^ Mao, Fangyuan; Li, Zhiyu; Wang, Zhili; Zhang, Chi; Rich, Thomas; Vickers-Rich, Patricia; Meng, Jin (2024-04-03). "Jurassic shuotheriids show earliest dental diversification of mammaliaforms". Nature. 628 (8008): 569–575. Bibcode:2024Natur.628..569M. doi:10.1038/s41586-024-07258-7. ISSN 0028-0836. PMID 38570681.

Mammalia

Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Cynodontia
Clade: Mammaliaformes

Synapsida

- see Synapsida

Cynodontia

see Cynodontia
Mammalia	see below↓

Mammalia

Australosphenida?

†Ambondro †Tendagurutherium? Monotremata?
†Henosferidae	Asfaltomylos Henosferus
†Ausktribosphenidae	Ausktribosphenos Bishops? Kryoparvus?

Monotremata

†Kollikodontidae?

†Steropodontidae

Parvopalus?
Steropodon

†Teinolophidae

Stirtodon?
Teinolophos

Ornithorhynchoidea

†Opalios
Ornithorhynchidae	†Dharragarra? †Monotrematum? †Obdurodon Ornithorhynchus †Patagorhynchus?
Tachyglossidae	†Megalibgwilia †Murrayglossus Tachyglossus Zaglossus

†Allotheria?

- see Allotheria
- includes multituberculates

Theriimorpha

- see below↓

Theriimorpha

†Gobiconodontidae

†Jeholodentidae

Jeholodens
Yanoconodon

†Klameliidae?

Ferganodon
Klamelia

Monotremata?
†Zhangheotheriidae	Anebodon Kiyatherium Maotherium Ningchengodon Origolestes Zhangheotherium
†Spalacotheriidae	Akidolestes Aliaga Cifellitherium Heishanlestes Infernolestes Lactodens Shalbaatar Spalacolestes Spalacotheridium Spalacotherium Spalacotheroides Symmetrodontoides Symmetrolestes Yaverlestes
Cladotheria	see below↓

Cladotheria

†Dryolestida

Brancatherulum? Dorsetodon Drescheratherium Euthlastus Henkelotherium Paurodon Tathiodon
Dryolestidae	Achyrodon Amblotherium Anthracolestes Beckumia Crusafontia Dryolestes Groebertherium? Guimarotodus Hercynodon Krebsotherium Lakotalestes? Laolestes Minutolestes Phascolestes Portopinheirodon Treslagosodon

†Meridiolestida

Amarillodon Austrotriconodon Bondesius Casamiquelia Cronopio Groebertherium? Lakotalestes? Leonardus Necrolestes Paraungulatum Quirogatherium Solanutherium
Mesungulatoidea	Coloniatherium Mesungulatum Orretherium Peligrotherium Reigitherium

†Donodontidae

Prototribosphenida

†Abelodon
†Amphibetulimus
†Amphitherium
†Arguimus
†Brancatherulum?
†Chunnelodon
†Magnimus
†Minimus
†Mozomus
†Nanolestes
†Palaeoxonodon
†Vincelestes
Monotremata?

†Peramuridae

Tribosphenida /
Boreosphenida

Theria

†Patagomaia
Eutheria	see Eutheria includes placentals
Metatheria	see Metatheria includes marsupials

Other taxa

Incertae sedis	†Atlasodon †Chronoperates †Microderson †Thereuodon †Tingamarra Saint Bathans Mammal UA 8699
Other taxa	Prototheria Yinotheria Eutriconodonta Amphilestidae Theriiformes Pantotheria Symmetrodonta Eupantotheria Paurodontidae Amphitheriidae Marsupionta

Extant mammal orders

Kingdom Animalia
Phylum Chordata
Subphylum Vertebrata
(unranked) Amniota

Yinotheria

Australosphenida	Monotremata (platypuses and echidnas)

Theria

Metatheria
(Marsupial inclusive)

Ameridelphia	Paucituberculata (shrew opossums) Didelphimorphia (opossums)
Australidelphia	Microbiotheria (monitos del monte) Notoryctemorphia (marsupial moles) Dasyuromorphia (quolls and dunnarts) Peramelemorphia (bilbies and bandicoots) Diprotodontia (kangaroos, koalas, and relatives)

Eutheria
(Placental inclusive)

Atlantogenata

Xenarthra	Cingulata (armadillos) Pilosa (anteaters and sloths)
Afrotheria	Afrosoricida (tenrecs, golden moles, and otter shrews) Macroscelidea (elephant shrews) Tubulidentata (aardvarks) Hyracoidea (hyraxes) Proboscidea (elephants) Sirenia (dugongs and manatees)

Boreoeutheria

Laurasiatheria	Eulipotyphla (hedgehogs, shrews, moles and relatives) Chiroptera (bats) Pholidota (pangolins) Carnivora (dogs, cats and relatives) Perissodactyla (horses, zebras, donkeys, rhinoceroses and tapirs) Artiodactyla (pigs, camels, hippos, deer, buffalo, gazelles, giraffes, whales, dolphins and relatives)
Euarchontoglires	Rodentia (rats, mice, guinea pigs, squirrels, beavers, chinchillas, porcupines, capybaras and relatives) Lagomorpha (rabbits, hares and pikas) Scandentia (treeshrews) Dermoptera (colugos) Primates (lorises, lemurs, tarsiers, monkeys, apes (including humans) and relatives)

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