Epstein Files Full PDF

CLICK HERE
Technopedia Center
PMB University Brochure
Faculty of Engineering and Computer Science
S1 Informatics S1 Information Systems S1 Information Technology S1 Computer Engineering S1 Electrical Engineering S1 Civil Engineering

faculty of Economics and Business
S1 Management S1 Accountancy

Faculty of Letters and Educational Sciences
S1 English literature S1 English language education S1 Mathematics education S1 Sports Education
teknopedia

  • Registerasi
  • Brosur UTI
  • Kip Scholarship Information
  • Performance
Flag Counter
  1. World Encyclopedia
  2. Varanidae
Varanidae
From Wikipedia, the free encyclopedia
(Redirected from Varanid lizard)
Family of lizards

Varanids
Temporal range:
Late Cretaceous – Holocene, 80–0 Ma
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N
Male Komodo dragons (Varanus komodoensis) fighting, Indonesia
Fossil of Saniwa, an extinct varanid known from the Eocene of North America
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Suborder: Anguimorpha
Superfamily: Varanoidea
Family: Varanidae
Merrem, 1820
Genera
  • Varanus
  • †Aiolosaurus
  • †Archaeovaranus
  • †Ovoo
  • †Paravaranus
  • †Proplatynotia
  • †Saniwa
  • †Saniwides
  • †Telmasaurus

The Varanidae are a family of lizards in the superfamily Varanoidea and order Anguimorpha. The family, a group of carnivorous and frugivorous lizards,[1] includes the living genus Varanus and a number of extinct genera more closely related to Varanus than to the earless monitor lizard (Lanthanotus).[2] Varanus includes the Komodo dragon (the largest living lizard), crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and Chinese crocodile lizard.[3] The oldest members of the family are known from the Late Cretaceous of Mongolia.[2]

Taxonomy

[edit]

The Varanidae were defined (using morphological characteristics) by Estes, de Queiroz and Gauthier (1988) as the clade containing the most recent common ancestor of Lanthanotus and Varanus and all of its descendants.[4] A similar definition was formulated by Conrad et al. (2008) (also using morphological data), who defined the Varanidae as the clade containing Varanus varius, Lanthanotus borneensis, and all descendants of their last common ancestor.[5] Using one of these definitions leads to the inclusion of the earless monitor lizard (L. borneensis) in the family Varanidae.

Lee (1997) created a different definition of the Varanidae, defining them as the clade containing Varanus and all taxa more closely related to Varanus than to Lanthanotus;[6][7] this definition explicitly excludes the earless monitor lizard from the Varanidae. Whether L. borneensis is included in or excluded from the Varanidae depends on the author; for example, Vidal et al. (2012) classify the earless monitor lizard as a member of a separate family Lanthanotidae,[8] while Gauthier et al. (2012) classify it as a member of Varanidae.[9]

Genera

[edit]
Genera marked with † are extinct

Genera included in Varanidae according to Dong et al., 2022[2]

  • †Ovoo Norell, Gao, & Conrad, 2008[10] (Mongolia, Late Cretaceous)
  • †Aiolosaurus Gao and Norell, 2000[10] (Mongolia, Late Cretaceous)
  • †Cherminotus Borsuk-Bialynicka, 1984 (Mongolia, Late Cretaceous)
  • †Saniwides Borsuk-Bialynicka, 1984 (Mongolia, Late Cretaceous)
  • †Paravaranus Borsuk-Bialynicka, 1984 (Mongolia, Late Cretaceous)
  • †Proplatynotia Borsuk-Bialynicka, 1984 (Mongolia, Late Cretaceous)
  • †Telmasaurus Gilmore, 1943[10] (Mongolia, Late Cretaceous)
  • †Saniwa Leidy, 1870 (Europe, North America, Eocene)
  • †Archaeovaranus Dong et al., 2022 (China, Eocene)
  • Varanus Shaw, 1790

Phylogeny

[edit]

Below is a cladogram from Dong et al. 2022.[2]

Varanidae

†Ovoo gurvel

†Telmasaurus grangeri

†Aiolosaurus oriens

†Saniwides mongoliensis

†Paravaranus angustifrons

†Proplatynotia longirostrata

†Saniwa ensidens

†Archaeovaranus lii

Varanus

Biology

[edit]
Gray's monitor (Varanus olivaceus) is a tree-dwelling varanid from the Philippines that primarily feeds on fruit

Monitor lizards are reputed to be among the most intelligent lizards. Most species forage widely and have large home ranges,[11] and many have high stamina.[12] Although most species are carnivorous, three arboreal species in the Philippines (Varanus olivaceus, Varanus mabitang, and Varanus bitatawa) are primarily frugivores.[1][13] Among species of living varanids, the limbs show positive allometry, being larger in larger-bodied species, although the feet become smaller as compared with the lengths of the other limb segments.[14]

Varanids possess unidirectional pulmonary airflow, including air-sacs akin to those of birds.[15]

References

[edit]
  1. ^ a b Welton LJ, Siler CD, Bennett D, Diesmos A, Duya MR, Dugay R, et al. (October 2010). "A spectacular new Philippine monitor lizard reveals a hidden biogeographic boundary and a novel flagship species for conservation". Biology Letters. 6 (5): 654–658. doi:10.1098/rsbl.2010.0119. PMC 2936141. PMID 20375042.
  2. ^ a b c d Dong L, Wang YQ, Zhao Q, Vasilyan D, Wang Y, Evans SE (March 2022). "A new stem-varanid lizard (Reptilia, Squamata) from the early Eocene of China". Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences. 377 (1847) 20210041. doi:10.1098/rstb.2021.0041. PMC 8819366. PMID 35125002.
  3. ^ Fry BG, Vidal N, Norman JA, Vonk FJ, Scheib H, Ramjan SF, et al. (February 2006). "Early evolution of the venom system in lizards and snakes". Nature. 439 (7076): 584–588. Bibcode:2006Natur.439..584F. doi:10.1038/nature04328. PMID 16292255. S2CID 4386245.
  4. ^ de Queiroz K, Gauthier J (1988). "Phylogenetic Relationships within Squamata". In Estes RJ, Pregill GK (eds.). Phylogenetic Relationships of the Lizard Families: Essays Commemorating Charles L. Camp. Stanford University Press. p. 166. ISBN 978-0-8047-1435-8. OCLC 16646258.
  5. ^ Conrad J (2008). "Phylogeny and systematics of Squamata (Reptilia) based on morphology". Bulletin of the American Museum of Natural History. 310: 1–182. doi:10.1206/310.1. hdl:2246/5915. S2CID 85271610.
  6. ^ WARNER R (1946). "Pectoral girdles vs. Hyobranchia in the snake genera Liotyphlops and Anomalepis". Science. 103 (2686): 720–722. doi:10.1126/science.103.2686.720. PMID 20986455.
  7. ^ Lee MS (June 2005). "Molecular evidence and marine snake origins". Biology Letters. 1 (2): 227–230. doi:10.1098/rsbl.2004.0282. PMC 1626205. PMID 17148173.
  8. ^ Vidal N, Marin J, Sassi J, Battistuzzi FU, Donnellan S, Fitch AJ, et al. (October 2012). "Molecular evidence for an Asian origin of monitor lizards followed by Tertiary dispersals to Africa and Australasia". Biology Letters. 8 (5): 853–855. doi:10.1098/rsbl.2012.0460. PMC 3441001. PMID 22809723.
  9. ^ Gauthier JA, Kearney M, Maisano JA, Rieppel O, Behlke AD (2012). "Assembling the Squamate Tree of Life: Perspectives from the Phenotype and the Fossil Record". Bulletin of the Peabody Museum of Natural History. 53 (1): 3–308. doi:10.3374/014.053.0101. S2CID 86355757.
  10. ^ a b c Conrad JL, Balcarcel AM, Mehling CM (2012). "Earliest example of a giant monitor lizard (Varanus, Varanidae, Squamata)". PLOS ONE. 7 (8) e41767. Bibcode:2012PLoSO...741767C. doi:10.1371/journal.pone.0041767. PMC 3416840. PMID 22900001.
  11. ^ Perry G, Garland Jr T (2002). "Lizard home ranges revisited: effects of sex, body size, diet, habitat, and phylogeny" (PDF). Ecology. 83 (7): 1870–1885. doi:10.1890/0012-9658(2002)083[1870:LHRREO]2.0.CO;2.
  12. ^ Clemente CJ, Withers PC, Thompson GG (2009). "Metabolic rate and endurance capacity in Australian varanid lizards (Squamata; Varanidae; Varanus)". Biological Journal of the Linnean Society. 97 (3): 664–676. doi:10.1111/j.1095-8312.2009.01207.x.
  13. ^ Greene HW (1986). Diet and Arboreality in the Emerald Monitor, Varanus prasinus, with Comments on the Study of Adaptation. Chicago: Field Museum of Natural History. OCLC 14915452. OL 7155983M.
  14. ^ Christian A, Garland Jr T (1996). "Scaling of limb proportions in monitor lizards (Squamata: Varanidae)" (PDF). Journal of Herpetology. 30 (2): 219–230. doi:10.2307/1565513. JSTOR 1565513.
  15. ^ Unidirectional Airflow In The Lungs Of Birds, Crocs And Now Monitor Lizards

External links

[edit]
  • Varanidae Archived 2010-03-27 at the Wayback Machine
  • v
  • t
  • e
Extant families of Squamata
  • Kingdom: Animalia
  • Phylum: Chordata
  • Class: Reptilia
  • Order: Squamata
Dibamia
  • Dibamidae
Gekkota
Pygopodoidea
  • Diplodactylidae
  • Carphodactylidae
  • Pygopodidae
Gekkonoidea
  • Eublepharidae
  • Sphaerodactylidae
  • Gekkonidae
  • Phyllodactylidae
Scincoidea
  • Scincidae
  • Xantusiidae
  • Cordylidae
  • Gerrhosauridae
Laterata /
Lacertoidea
  • Lacertidae
Gymnophthalmoidea
  • Alopoglossidae
  • Gymnophthalmidae
  • Teiidae
Amphisbaenia
  • Amphisbaenidae
  • Bipedidae
  • Blanidae
  • Cadeidae
  • Rhineuridae
  • Trogonophidae
Anguimorpha
Paleoanguimorpha
  • Shinisauridae
  • Lanthanotidae
  • Varanidae
Neoanguimorpha
  • Helodermatidae
  • Xenosauridae
  • Anguidae
  • Anniellidae
  • Diploglossidae
Iguania
Acrodonta
  • Agamidae
  • Chamaeleonidae
Pleurodonta
  • Corytophanidae
  • Crotaphytidae
  • Dactyloidae
  • Hoplocercidae
  • Iguanidae
  • Leiocephalidae
  • Leiosauridae
  • Liolaemidae
  • Opluridae
  • Phrynosomatidae
  • Polychrotidae
  • Tropiduridae
Serpentes
Scolecophidia
  • Anomalepididae
  • Gerrhopilidae
  • Leptotyphlopidae
  • Typhlopidae
  • Xenotyphlopidae
Alethinophidia
  • Acrochordidae
  • Aniliidae
  • Anomochilidae
  • Atractaspididae
  • Boidae
  • Bolyeriidae
  • Colubridae
  • Cylindrophiidae
  • Elapidae
  • Loxocemidae
  • Pythonidae
  • Tropidophiidae
  • Uropeltidae
  • Viperidae
  • Xenopeltidae
  • v
  • t
  • e
Varanoidea
  • Kingdom: Animalia
  • Phylum: Chordata
  • Class: Reptilia
  • Order: Squamata
Extant species
Helodermatidae
  • Chiapan beaded lizard (H. alvarezi)
  • Guatemalan beaded lizard (H. charlesbogerti)
  • Rio Fuerte beaded lizard (H. exasperatum)
  • Mexican beaded lizard (H. horridum)
  • Gila monster (H. suspectum)
Lanthanotus
  • Earless monitor lizard (L. borneensis)
Varanus
  • Asian water monitor (V. salvator)
  • Black tree monitor (V. beccarii)
  • Black-headed monitor (V. tristis)
  • Bengal monitor (V. bengalensis)
  • Black-spotted ridge-tailed monitor (V. baritji)
  • Blue-tailed monitor (V. doreanus)
  • Crocodile monitor (V. salvadorii)
  • Desert monitor (V. griseus)
  • Dumeril's monitor (V. dumerilii)
  • Emerald tree monitor (V. prasinus)
  • Gray's monitor (V. olivaceus)
  • Kimberely rock monitor (V. glauerti)
  • Komodo dragon (V. komodoensis)
  • Lace monitor (V. varius)
  • Mangrove monitor (V. indicus)
  • Merten's water monitor (V. mertensi)
  • Mitchell's water monitor (V. mitchelli)
  • Nile monitor (V. niloticus)
  • Northern Sierra Madre forest monitor (V. bitatawa)
  • Peacock monitor (V. auffenbergi)
  • Peach-throated monitor (V. jobiensis)
  • Perentie (V. giganteus)
  • Pygmy mulga monitor (V. gilleni)
  • Quince monitor (V. melinus)
  • Rock monitor (V. albigularis)
  • Pilbara monitor (V. bushi)
  • Rennel Island monitor (V. juxtindicus)
  • Rosenberg's monitor (V. rosenbergi)
  • Roughneck monitor (V. rudicollis)
  • Short-tailed pygmy monitor (V. brevicauda)
  • Spiny-tailed monitor (V. acanthurus)
  • Turquoise monitor (V. caerulivirens)
  • Sand goanna (V. gouldii)
  • Savannah monitor (V. exanthematicus)
  • Yellow monitor (V. flavescens)
  • Yellow-spotted monitor (V. panoptes)
  • Yemen monitor (V. yemenensis)
Related categories
  • Varanidae
  • Cretaceous lizards
  • Helodermatidae
Fossil taxa
†Palaeovaranidae
  • Palaeovaranus
  • Eosaniwa
  • Paranecrosaurus
Varanidae
  • Megalania (Varanus priscus)
  • Saniwa
  • Varanus amnhophilis
Others
  • Arcanosaurus
  • Aiolosaurus
  • Cherminotus
  • Gobiderma
  • Morohasaurus
  • Ovoo
  • Telmasaurus
  • Palaeosaniwa
  • Saniwides
  • Estesia
Taxon identifiers
Varanidae
  • Wikidata: Q3082684
  • Wikispecies: Varanidae
  • AFD: Varanidae
  • BOLD: 1099
  • CoL: 6273W
  • EoL: 8115
  • EPPO: 1VARAF
  • GBIF: 5023
  • iNaturalist: 39392
  • IRMNG: 103447
  • ITIS: 174114
  • NCBI: 8555
  • Open Tree of Life: 713776
  • Paleobiology Database: 38035
  • Plazi: 592C4D59-0A61-FFF1-2BC0-3E8E5D32B4E3
  • WoRMS: 395082
Authority control databases Edit this at Wikidata
National
  • Czech Republic
Other
  • Yale LUX
Retrieved from "https://en.wikipedia.org/w/index.php?title=Varanidae&oldid=1314555801"
Categories:
  • Varanidae
  • Lizard families
  • Taxa named by Blasius Merrem
  • Extant Campanian first appearances
Hidden categories:
  • Articles with short description
  • Short description is different from Wikidata
  • Articles with 'species' microformats
  • Webarchive template wayback links

  • indonesia
  • Polski
  • العربية
  • Deutsch
  • English
  • Español
  • Français
  • Italiano
  • مصرى
  • Nederlands
  • 日本語
  • Português
  • Sinugboanong Binisaya
  • Svenska
  • Українська
  • Tiếng Việt
  • Winaray
  • 中文
  • Русский
Sunting pranala
url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url url
Pusat Layanan

UNIVERSITAS TEKNOKRAT INDONESIA | ASEAN's Best Private University
Jl. ZA. Pagar Alam No.9 -11, Labuhan Ratu, Kec. Kedaton, Kota Bandar Lampung, Lampung 35132
Phone: (0721) 702022
Email: pmb@teknokrat.ac.id