Chuandongocoelurus

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Chuandongocoelurus - Wikipedia

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Chuandongocoelurus Temporal range: Middle Jurassic, 165 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓

Skeletal diagram showing known remains
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Clade:	Theropoda
Genus:	†Chuandongocoelurus He, 1984
Species:	†C. primitivus
Binomial name
†Chuandongocoelurus primitivus He, 1984

Chuandongocoelurus (/tʃwɑːnˌdɒŋəsɪˈljʊərəs/ chwahn-DONG-ə-si-LURE-əs) is a genus of carnivorous tetanuran theropod dinosaur from the Jurassic of China.

Discovery and naming

The type species Chuandongocoelurus primitivus was first described and named by Chinese paleontologist He Xinlu in 1984. The generic name combines references to the Chuandong in Sichuan Province and the theropod genus Coelurus, itself named after the Greek κοῖλος, koilos, meaning "hollow" and οὐρά, oura, meaning "tail". The specific name means "the primitive one" in Latin, a reference to the great age of the find.^[1]

He assigned two partial skeletons to Chuandongocoelurus. The holotype, a thighbone, is part of specimen CCG 20010. Vertebrae, pelvic bones and hindlimb elements, also catalogued under this inventory number, may belong to the same individual. The specimen has unfused neurocentral sutures in its vertebrae, meaning that the animal was immature at the time of death. The second specimen, CCG 20011, is a set of neck vertebrae from a much larger individual.^[1] In 2012, it was concluded that both specimens represent different taxa, probably not even closely related. CCG 20011 was found to instead share similarities with the ceratosaur Elaphrosaurus.^[2] The formation in which it was discovered was the Lower Shaximiao Formation, meaning Chuandongocoelurus dates to the Bathonian or Callovian stage of the Middle Jurassic.

The holotype thighbone has a length of 201 millimetres.^[1]

Classification

He placed Chuandongocoelurus in the Coeluridae,^[1] at the time a wastebasket taxon including almost all small theropods. David Bruce Norman in 1990 considered it to be an indeterminate theropod.^[3] More recently, Roger Benson (2008, 2010) and Benson et al. (2010) found it to be the sister taxon of Monolophosaurus, together forming a clade belonging either to Megalosauroidea^[4]^[5] or outside of Megalosauroidea in the Tetanurae.^[6] In 2012, Matthew Carrano et al. found Chuandongocoelurus outside of the Megalosauroidea.^[2]

The cladogram below follows Rauhut and Pol (2019):^[7]

Averostra

Ceratosauria

Tetanurae

Chuandongocoelurus

References

^ ^a ^b ^c ^d He, 1984. The vertebrate fossils of Sichuan. Sichuan Scientific and Technological Publishing House. 168 pp.
^ ^a ^b M.T. Carrano, R.B.J. Benson, and S.D. Sampson, 2012, "The phylogeny of Tetanurae (Dinosauria: Theropoda)", Journal of Systematic Palaeontology 10(2): 211-300
^ Norman, David B. (1990). Problematic Theropoda: "Coelurosaurs". p. 280-305 in David B. Weishampel, et al. (eds.), The Dinosauria. University of California Press, Berkeley, Los Angeles, Oxford.
^ Benson, 2008. A new theropod phylogeny focussing on basal tetanurans, and its implications for European 'megalosaurs' and Middle Jurassic dinosaur endemism. Journal of Vertebrate Paleontology. 51A
^ Benson, R.B.J. (2010). "A description of Megalosaurus bucklandii (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods". Zoological Journal of the Linnean Society. 158 (4): 882–935. doi:10.1111/j.1096-3642.2009.00569.x.
^ Benson, Brusatte and Carrano, 2010. A new clade of large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic. Naturwissenschaften. 97, 71-78
^ Rauhut, Oliver W. M.; Pol, Diego (2019-12-11). "Probable basal allosauroid from the early Middle Jurassic Cañadón Asfalto Formation of Argentina highlights phylogenetic uncertainty in tetanuran theropod dinosaurs". Scientific Reports. 9 (1): 18826. doi:10.1038/s41598-019-53672-7. ISSN 2045-2322. PMC 6906444. PMID 31827108.

External links

Details on Chuandongocoelurus, from the Dinosaur Mailing List (includes full redescription)

Theropoda

Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Clade: Avemetatarsalia
Clade: Dinosauria

Avemetatarsalia

see Avemetatarsalia
Theropoda	see below↓

Theropoda

Neotheropoda

Coelophysoidea

Dracoraptor? Panguraptor Pendraig Podokesaurus Powellvenator
Coelophysidae	Camposaurus Coelophysis Lucianovenator Megapnosaurus? Panguraptor? Procompsognathus Pterospondylus? Segisaurus

Averostra

- see below↓

Dubious neotheropods

Teinurosaurus
Halticosaurus

Averostra

Dornraptor
Lophostropheus?

Ceratosauria

Berberosaurus Fosterovenator? Jubbulpuria Lukousaurus? Saltriovenator
Ceratosauridae	Ceratosaurus Genyodectes

Abelisauroidea

Noasauridae

Afromimus? Austrocheirus? Berthasaura Compsosuchus? Dahalokely? Deltadromeus? Genusaurus? Kiyacursor Ligabueino Spinostropheus
Elaphrosaurinae	Elaphrosaurus Huinculsaurus Limusaurus
Noasaurinae	Laevisuchus Masiakasaurus Noasaurus Velocisaurus Vespersaurus

Abelisauridae

Majungasaurinae

Carnotaurinae

Majungasaurinae?

Brachyrostra

Dahalokely? Ekrixinatosaurus Elemgasem Guemesia Ilokelesia Llukalkan Niebla Rahiolisaurus? Skorpiovenator Thanos
Furileusauria	Abelisaurus Aucasaurus Caletodraco Carnotaurus Elemgasem? Koleken Pycnonemosaurus Quilmesaurus Viavenator

Tetanurae	see below↓

Tetanurae

Orionides

Megalosauroidea

Yunyangosaurus
Piatnitzkysauridae	Condorraptor Marshosaurus Piatnitzkysaurus Xuanhanosaurus?

Megalosauria

Streptospondylus?

Megalosauridae

Eustreptospondylus
Megalosaurinae	Duriavenator Megalosaurus Torvosaurus Wiehenvenator
Afrovenatorinae	Afrovenator Dubreuillosaurus Leshansaurus Magnosaurus Piveteausaurus Poekilopleuron?

Spinosauridae

Baryonychinae

Baryonyx Protathlitis Riojavenatrix Suchosaurus? Vallibonavenatrix?
Ceratosuchopsini	Ceratosuchops Riparovenator Suchomimus

Spinosaurinae

Camarillasaurus? Ichthyovenator Irritator Oxalaia Siamosaurus
Spinosaurini	Sigilmassasaurus Spinosaurus

Avetheropoda	see below↓

Avetheropoda

Gasosaurus?
Lourinhanosaurus?

Carnosauria

Allosauroidea

Metriacanthosauridae

Xuanhanosaurus? Yangchuanosaurus
Metriacanthosaurinae	Alpkarakush Metriacanthosaurus Shidaisaurus Siamotyrannus Sinraptor

Carcharodontosauridae

Carcharodontosaurinae

Carcharodontosaurus
Giganotosaurini	Giganotosaurus Mapusaurus Meraxes Taurovenator Tyrannotitan

Megaraptora?

Aoniraptor Bahariasaurus? Deltadromeus? Fukuiraptor Phuwiangvenator Rapator Siats? Chilantaisaurus?
Megaraptoridae	Aerosteon Australovenator Maip Megaraptor Murusraptor Orkoraptor Tratayenia

Coelurosauria

- see below↓

Coelurosauria

Tyrannoraptora

Tyrannosauroidea

Chingkankousaurus Labocania?
Coeluridae?	Coelurus Tanycolagreus?
Proceratosauridae	Guanlong Kileskus Proceratosaurus Sinotyrannus Yutyrannus

Albertosaurus
Gorgosaurus

Tyrannosaurinae

Asiatyrannus Nanuqsaurus
Alioramini	Alioramus Qianzhousaurus
Daspletosaurini	Daspletosaurus Thanatotheristes
Teratophoneini	Bistahieversor? Dynamoterror Labocania? Lythronax Teratophoneus
Tyrannosaurini	Tarbosaurus Tyrannosaurus Zhuchengtyrannus

Maniraptoromorpha	see below↓

Dubious coelurosaurs	Iliosuchus Kakuru Phaedrolosaurus Shanyangosaurus

Maniraptoromorpha

Neocoelurosauria

Aniksosaurus
Compsognathidae	Aniksosaurus? Aristosuchus Beipiaognathus? Compsognathus Huaxiagnathus Mirischia Scipionyx? Sinocalliopteryx Sinosauropteryx Xunmenglong

Maniraptoriformes

Compsognathidae?

Ornithomimosauria

Macrocheiriformes

Harpymimus Pelecanimimus Shenzhousaurus
Deinocheiridae	Beishanlong Deinocheirus Garudimimus Harpymimus? Paraxenisaurus? Tyrannomimus
Ornithomimidae	Aepyornithomimus Anserimimus Archaeornithomimus "Coelosaurus" Dromiceiomimus Gallimimus Ornithomimus Qiupalong Rativates Sinornithomimus Struthiomimus Tototlmimus

Maniraptora

- see Maniraptora
- includes birds

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